AI Nagahama¹, Shuichiro Tagane², Nguyen Van Ngoc³, Hoang Thi Binh³, Truong Quang Cuong⁴, Nguyen Thi Anh Thu³, Kojin Tsuchiya⁵, Meng Zhang¹, Hoang Viet Thanh³, Nguyen Cuong Thinh³, Yoshihisa Suyama⁶, Ayumi Matsuo⁶, Shun K. Hirota⁶, Etsuko Moritsuka¹, Hironori Toyama⁷, Hidetoshi Nagamasu⁸, Naiki Akiyo⁷ and Tetsukazu Yahara¹, (1)Kyushu University, Fukuoka, Japan, (2)The Kagoshima University Museum, Kagoshima, Japan, (3)Dalat University, Dalat, Vietnam, (4)Bidoup-Nui Ba National Park, Dalat, Vietnam, (5)The Univeristy of Tokyo, Tokyo, Japan, (6)Tohoku University, Osaki, Japan, (7)University of the Ryukyus, Taketomi, Japan, (8)The Kyoto University Museum, Kyoto, Japan