Timothy Peter Burt, University of Durham, Durham, United Kingdom, Fred Worrall, Durham University, Durham, United Kingdom, Nicholas J K Howden, University of Bristol, Bristol, BS8, United Kingdom, Helen P Jarvie, Centre for Ecology and Hydrology, Wallingford, Oxfordshire, United Kingdom, Andrew D Pratt, Field Studies Council, Slapton Ley Field Centre, Slapton, United Kingdom and Thomas Hutchinson, Plymouth University, Plymouth, United Kingdom