Xuehe Lu¹, Weimin Ju¹, Jing Li¹, Holly Croft^2,3, Jingming Chen³ and Yiqi Luo⁴, (1)Nanjing University, Nanjing, China, (2)University of Sheffield, Sheffield, United Kingdom, (3)University of Toronto, Toronto, Canada, (4)Department of Biological Sciences, Northern Arizona University, Flagstaff, AZ, United States