Haiqiang Gao^1,2, Shuguang Liu³, Weizhi LU⁴, Andrew R Smith⁵, Juyang Liao⁶, Zhenming Wu⁷, Li Xiao⁸, Zhao Wang⁸, Xi Peng⁸, Qinyuan Li⁸ and Yujun Feng⁸, (1)Changsha, Hunan, China, (2)Central South University of Forestry and Technology, National Engineering Laboratory for Applied Technology of Forestry & Ecology in South China, Changsha, China, (3)Central South University of Forestry and Technology, National Engineering Laboratory of Applied Technology of Forestry & Ecology for Southern China, Changsha, China, (4)State Oceanic Administration, Dalian, China, (5)Bangor University, Bangor, United Kingdom, (6)Hunan Forest Botanical Garden, Changsha, China, (7)Forestry Department of Hunan Province, Changsha, China, (8)Central South University of Forestry and Technology, Changsha, China