Yizhao Chen¹, Annemarie H. Eckes¹, Tim Tito Rademacher^2,3, Andrew D Richardson⁴, Patrick Fonti⁵, Jim Le Moine⁶ and Andrew D Friend¹, (1)University of Cambridge, Cambridge, United Kingdom, (2)Harvard University, Department of Organismic and Evolutionary Biology, Cambridge, United States, (3)Northern Arizona University, Center for Ecosystem Science and Society, Flagstaff, United States, (4)Northern Arizona University, Flagstaff, AZ, United States, (5)Swiss Federal Research Institute WSL, Birmensdorf, Switzerland, (6)Northern Arizona University, Flagstaff, United States