William Cox, University of Guelph, Integrative Biology, Guelph, ON, Canada, Merritt R Turetsky, University of Guelph, Guelph, ON, Canada, Thomas A Douglas, USA CRREL Alaska, Fort Wainwright, AK, United States, Catherine M Dieleman, University of Guelph, Department of Integrative Biology, Guelph, ON, Canada and Evan S Kane, Michigan Technological University, School of Forest Resources and Environmental Sciences, Houghton, MI, United States