Zhenxiu JIA¹, Shuwen Pei², Alfonso Benito-Calvo³, Dongdong Ma², Laura Sanchez-Romero³ and Qi Wei², (1)ITP Institute of Tibetan Plateau Research, Chinese Academy of Sciences, Beijing, China, (2)Key Laboratory of Vertebrate Evolution and Human Origins, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing, China, (3)Centro Nacional de Investigación sobre la Evolución Humana (CENIEH), Burgos, Spain