Jongkuk Choi¹, Robert J W Brewin², Jae Hoon Noh¹ and Xuerong Sun³, (1)Korea Institute of Ocean Science and Technology, Busan, South Korea, (2)College of Life and Environmental Sciences, University of Exeter, Penryn, United Kingdom, (3)State Key Laboratory of Estuarine and Coastal Research, East China Normal University, Shanghai, China