Tsuyoshi Watanabe¹, Satoshi Nagai², Jun Kikuchi³, Nobuharu Inaba², Yukiko Taniuchi⁴, Taiga Asakura³, Hiroaki Kurogi⁵, Seinen Chow², Tsutomu Tomoda⁶, Daisuke Ambe⁷ and Daisuke Hasegawa¹, (1)Tohoku National Fisheries Research Institute, Japan Fisheries Research and Education Agency, Shiogama, Japan, (2)National Research Institute of Fisheries Science, Japan Fisheries Research and Education Agency, Japan, (3)RIKEN Center for Sustainable Resource Science, Japan, (4)Hokkaido National Fisheries Research Institute, Japan Fisheries Research and Education Agency, Kushiro, Japan, (5)National Research Institute of Fisheries Science, Japan Fisheries Research and Education Agency, Yokosuka, Japan, (6)National Research Institute of Aquaculture, Japan Fisheries Research and Education Agency, Japan, (7)National Research Institute of Fisheries Science, Japan Fisheries Research and Education Agency, Yokohama, Japan