Leon Dlugosch¹, Anja Poehlein², Bernd Wemheuer², Thomas H. Badewien³, Rolf Daniel² and Meinhard Simon¹, (1)University of Oldenburg, Institute for Chemistry and Biology of the Marine Environment (ICBM), Biology of Geological Processes - Aquatic Microbial Ecology, Oldenburg, Germany, (2)University of Göttingen, Institut für Mikrobiologie und Genetik, Göttingen, Germany, (3)University of Oldenburg, Institute for Chemistry and Biology of the Marine Environment, Wilhelmshaven, Germany