Beom-Sik Kim¹, Chang-Keun Kang², Hae Kun Jung³, Hyun-Woo Kim⁴, Hyun Je Park⁵, Jongwon Park¹ and Chung Il Lee⁵, (1)Gangneung-Wonju National University, Department of Marine Ecology and Environment, Gangneung-si, South Korea, (2)Gwangju Institute of Science and Technology, School of Environment and Energy Engineering, Gwangju, South Korea, (3)National Institute of Fisheries Science, Ocean Climate and Ecology Research Division, Busan, South Korea, (4)Pukyong National University, Department of Marine Biology, Busan, South Korea, (5)Gangneung-Wonju National University, Department of Marine Ecology and Environment, Gangneung, South Korea