Sachiko Horii, Nagasaki Field Station, Japan Fisheries Research and Education Agency, Fisheries Resources Institute, Nagasaki, Japan, Hiroaki Saito, The University of Tokyo, Atmosphere and Ocean Research Institute, Kashiwa, Japan, Keitaro Fukushima, Fukushima University, Faculty of Food and Agricultural Sciences, Fukushima, Japan, Yuji Onishi, Research Institute for Humanity and Nature, Kyoto, Japan, Keisuke Koba, Center for Ecological Research, Kyoto University, Kyoto, Japan, Ken Furuya, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan and Kazutaka Takahashi, The University of Tokyo, Graduate School of Agricultural and Life Sciences, Tokyo, Japan