Kota Ishikawa^1,2, Takuma Masui³, Keita Koeda⁴ and Michio Kondoh^1,2, (1)Tohoku University, Graduate School of Life Sciences, Sendai, Japan, (2)Tohoku University & JAMSTEC Advanced Institute for Marine Ecosystem Change, Sendai, Japan, (3)University of the Ryukyus, Graduate School of Science and Engineering, Okinawa, Japan, (4)University of the Ryukyus, Faculty of Science, Okinawa, Japan