Zhen Wu¹, Ioannis Tsakalakis², John Casey³, Michael J Follows³ and Joseph John Vallino⁴, (1)Peking University, Beijing, China, (2)Massachusetts Institute of Technology, Cambridge, United States, (3)Massachusetts Institute of Technology, Department of Earth, Atmospheric and Planetary Sciences, Cambridge, United States, (4)Marine Biological Laboratory, Ecosystems Center, Woods Hole, MA, United States