Takehiro Shimonaka¹, Shigeyoshi Otosaka², Takafumi Kataoka³, Junya Hirai², Taku Wagawa⁴, Misato Nakae⁴, Kay Sakuma⁴, Kazutaka Takahashi¹ and Taketoshi Kodama¹, (1)The University of Tokyo, Graduate School of Agricultural and Life Sciences, Tokyo, Japan, (2)The University of Tokyo, Atmosphere and Ocean Research Institute, Kashiwa, Japan, (3)Fukui Prefectural University, Faculty of Marine Science and Technology, Obama, Japan, (4)Japan Fisheries Research and Education Agency, Fisheries Resources Institute, Niigata, Japan