Tomohiro Nakamura, Hokkaido University, Institute of Low Temperature Science, PORC, Sapporo, Japan, Yusuke Takahashi, Hokkaido University, Graduate School of Environmental Science, Sapporo, Japan, Takuya Nakanowatari, Japan Fisheries Research and Education Agency, Fisheries Resources Institute, Kushiro, Japan and Humio Mitsudera, Institute of Low Temperature Science, Hokkaido University, Sapporo, Japan