Amy Pickard¹, Alanna Grant², Nathan Callaghan³, Chris Evans³, Anna Belcher⁴, Rebecca McKenzie⁴, Bryan M. M. Spears⁴ and Science team of the NERC BIOPOLE project, (1)UK Centre for Ecology & Hydrology, Penicuik, UNITED KINGDOM, (2)UK Centre for Ecology & Hydrology, Edinburgh, United Kingdom, (3)UK Centre for Ecology & Hydrology, Bangor, United Kingdom, (4)UK Centre for Ecology & Hydrology, Penicuik, United Kingdom