Jiaxing Liu, South China Sea Institute of Oceanology, Chinese Academy of Sciences No.164 West Xingang Road, Guangzhou, China, CHINA, Yehui Tan, SCSIO South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou, China and Hongbin Liu, HKUST, Kowloon, Hong Kong; Hong Kong University of Science and Technology, Division of Life Science, Hong Kong, Hong Kong