Jiaxin Chen¹, Yu Wang², Yibin Huang², Qi Chen³, Chenwen Xue², Jinxin Xu², Xiaomeng Wang⁴, Chen He⁵, Shi Quan⁶, Helena Osterholz⁷, Christian Lønborg⁸, Yu Cong², Nianzhi Jiao⁹ and Qiang Zheng¹⁰, (1)State Key Laboratory for Marine Environmental Science, College of Ocean and Earth Sciences, Xiamen University, Xiamen, China, (2)Xiamen University, Xiamen, China, (3)Oregon State University, Department of Microbiology, Corvallis, United States, (4)Southern University of Science and Technology, Shenzhen, China, (5)China University of Petroleum, Beijing, China, (6)China University of Petroleum, State Key Laboratory of Heavy Oil Processing, Beijing, China, (7)Leibniz Institute for Baltic Sea Research, Marine Chemistry, Rostock, Germany, (8)Aarhus University, Roskilde, Denmark, (9)Xiamen University, Innovation Research Center for Carbon Neutralization, Fujian Key Laboratory of Marine Carbon Sequestration, Xiamen, China, (10)State Key Laboratory of Marine Environmental Science, Xiamen University, Xiamen, China