Robert E Todd, Woods Hole Oceanographic Institution, Woods Hole, MA, United States, Yuichiro Takeshita, Monterey Bay Aquarium Research Institute, Moss Landing, CA, United States, Daniel L Rudnick, Scripps Institution of Oceanography, La Jolla, CA, United States, Christopher A Edwards, University of California Santa Cruz, Santa Cruz, CA, United States and Jann Paul Mattern, University of California Santa Cruz, Ocean Sciences Department, Santa Cruz, United States; Dalhousie University, Halifax, NS, Canada